In dense underwater sensor networks (UWSN), the major confronts are high error probability, incessant variation in topology of sensor nodes, and much energy consumption for data transmission. However, there are some remarkable applications of UWSN such as management of seabed and oil reservoirs, exploration of deep sea situation and prevention of aqueous disasters. In order to accomplish these applications, ignorance of the limitations of acoustic communications such as high delay and low bandwidth is not feasible. In this paper, we propose Adaptive mobility of Courier nodes in Threshold-optimized Depth-based routing (AMCTD), exploring the proficient amendments in depth threshold and implementing the optimal weight function to achieve longer network lifetime. We segregate our scheme in 3 major phases of weight updating, depth threshold variation and adaptive mobility of courier nodes. During data forwarding, we provide the framework for alterations in threshold to cope with the sparse condition of network. We ultimately perform detailed simulations to scrutinize the performance of our proposed scheme and its comparison with other two notable routing protocols in term of network lifetime and other essential parameters. The simulations results verify that our scheme performs better than the other techniques and near to optimal in the field of UWSN.