We consider a multi-hop distributed hypothesis testing problem with multiple decision centers (DCs) for testing against independence and where the observations obey some Markov chain. For this system, we characterize the fundamental type-II error exponents region, i.e., the type-II error exponents that the various DCs can achieve simultaneously, under expected rate-constraints. Our results show that this fundamental exponents region is boosted compared to the region under maximum-rate constraints, and that it depends on the permissible type-I error probabilities. When all DCs have equal permissible type-I error probabilities, the exponents region is rectangular and all DCs can simultaneously achieve their optimal type-II error exponents. When the DCs have different permissible type-I error probabilities, a tradeoff between the type-II error exponents at the different DCs arises. New achievability and converse proofs are presented. For the achievability, a new multiplexing and rate-sharing strategy is proposed. The converse proof is based on applying different change of measure arguments in parallel and on proving asymptotic Markov chains. For the special cases $K = 2$ and $K = 3$, we provide simplified expressions for the exponents region; a similar simplification is conjectured for arbitrary $K\geq 2$.