A marked lattice is a $d$-dimensional Euclidean lattice, where each lattice point is assigned a mark via a given random field on ${\mathbb Z}^d$. We prove that, if the field is strongly mixing with a faster-than-logarithmic rate, then for every given lattice and almost every marking, large spheres become equidistributed in the space of marked lattices. A key aspect of our study is that the space of marked lattices is not a homogeneous space, but rather a non-trivial fiber bundle over such a space. As an application, we prove that the free path length in a crystal with random defects has a limiting distribution in the Boltzmann-Grad limit.