Eulerian polynomials are fundamental in combinatorics and algebra. In this paper we study the linear transformation $\mathcal{A} : \mathbb{R}[t] \to \mathbb{R}[t]$ defined by $\mathcal{A}(t^n) = A_n(t)$, where $A_n(t)$ denotes the $n$-th Eulerian polynomial. We give combinatorial, topological and Ehrhart theoretic interpretations of the operator $\mathcal{A}$, and investigate questions of unimodality and real-rootedness. In particular, we disprove a conjecture by Brenti (1989) concerning the preservation of real zeros, and generalize and strengthen recent results of Haglund and Zhang (2019) on binomial Eulerian polynomials.