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Complement Submodular Information Measures for Balanced and Robust Data Selection A Proof of a Conjecture on Positive and Negative Square Energies of Unicyclic Graphs Laplacian Spectrum of the Weakly Zero-Divisor Graph of a Finite Commutative Ring An identity for second Eulerian numbers via lattice-point counting $t$-tone edge coloring of graphs Constructing Maximal Bumpless Pipedreams for Double Grothendieck Polynomials Mubayi's Polynomial-Ideal Conjecture and Cover-Ideal Turán Methods Implicit Binarization via Complex Phase Dynamics in Combinatorial Optimization The limits of Schur multipliers in Pólya conversion problems for the $q$-permanent function Universality theorems for generalized splines Framing Triangulations for Arbitrary Integer Flow Polytopes On the Common Generalization of Gentle Algebras and Framed Directed Acyclic Graphs The complexity of frugal digraph homomorphisms Chaotic and periodic behavior of jeu de taquin on infinite Young tableaux Enumerating Pattern Avoiding Parking Functions Incidence toric ideals and three-point functions Unique Winning Opening Move in Three-Row Chomp Strong majority colorings of graphs A Balancing Theorem for Spanning Trees of Rectangular Grid Graphs Spectral radius and edge-disjoint connected factors of graphs New invariants for rank metric codes, with applications to the classification of rank two semifields of order 256 Flexible DP-4-coloring of planar graphs without 4-cycles and intersecting triangles Balanced intersection size distributions in projective planes List Reconstruction Problem with List Size Two Is Dimensionality a Barrier for Retrieval Models? The INIEP: Irreducible and Positive Realizations The number of Pfaffian orientations on punctured polygonally cellulated surfaces Explicit Construction of Polytopes whose Ehrhart Polynomials Realize any Given Sign Pattern Finite-state enumeration of adjacency-constrained 132-avoiding permutations AMDS and quantum AMDS Constacyclic codes of length $4p^ς$ over $\mathbb{F}_{{p}^{m}}$
Revisiting Shao and Sokal's $B_2$ index of phylogenetic b...
François Bienvenu, Gabriel Cardona, Celine Scornavacca · 2020-10-16 · via math.CO updates on arXiv.org

Measures of phylogenetic balance, such as the Colless and Sackin indices, play an important role in phylogenetics. Unfortunately, these indices are specifically designed for phylogenetic trees, and do not extend naturally to phylogenetic networks (which are increasingly used to describe reticulate evolution). This led us to consider a lesser-known balance index, whose definition is based on a probabilistic interpretation that is equally applicable to trees and to networks. This index, known as the $B_2$ index, was first proposed by Shao and Sokal in 1990. Surprisingly, it does not seem to have been studied mathematically since. Likewise, it is used only sporadically in the biological literature, where it tends to be viewed as arcane. In this paper, we study mathematical properties of $B_2$ such as its expectation and variance under the most common models of random trees and its extremal values over various classes of phylogenetic networks. We also assess its relevance in biological applications, and find it to be comparable to that of the Colless and Sackin indices. Altogether, our results call for a reevaluation of the status of this somewhat forgotten measure of phylogenetic balance.