Ostrom and Wagner (1959) proved that if the automorphism group $G$ of a finite projective plane $π$ acts $2$-transitively on the points of $π$, then $π$ is isomorphic to the Desarguesian projective plane and $G$ is isomorphic to $\mathrm{PΓL}(3,q)$ (for some prime-power $q$). In the more general case of a finite rank $2$ irreducible spherical building, also known as a \emph{generalized polygon}, the theorem of Fong and Seitz (1973) gave a classification of the \emph{Moufang} examples. A conjecture of Kantor, made in print in 1991, says that there are only two non-classical examples of flag-transitive generalized quadrangles up to duality. Recently, the authors made progress toward this conjecture by classifying those finite generalized quadrangles which have an automorphism group $G$ acting transitively on antiflags. In this paper, we take this classification much further by weakening the hypothesis to $G$ being transitive on ordered pairs of collinear points and ordered pairs of concurrent lines.