We prove a `motivic' analogue of the Weyl character formula, computing the Euler characteristic of a line bundle on a generalized flag manifold $G/B$ multiplied either by a motivic Chern class of a Schubert cell, or a Segre analogue of it. The result, given in terms of Demazure-Lusztig (D-L) operators, recovers formulas found by Brubaker, Bump and Licata for the Iwahori-Whittaker functions of the principal series representation of a $p$-adic group. In particular, we obtain a new proof of the classical Casselman-Shalika formula for the spherical Whittaker function. The proofs are based on localization in equivariant K theory, and require a geometric interpretation of how the Hecke dual (or inverse) of a D-L operator acts on the class of a point. We prove that the Hecke dual operators give Grothendieck-Serre dual classes of the motivic classes, a result which might be of independent interest. In an Appendix joint with Dave Anderson we show that if the line bundle is trivial, we recover a generalization of a classical formula by Kostant, Macdonald, Shapiro and Steinberg for the Poincar{é} polynomial of $G/B$; the generalization we consider is due to Akyıldız and Carrell and replaces $G/B$ by any smooth Schubert variety.