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MPCS: Neuroplastic Continual Learning via Multi-Component Plasticity and Topology-Aware EWC Combining Trained Models in Reinforcement Learning Training Non-Differentiable Networks via Optimal Transport ShiftLIF: Efficient Multi-Level Spiking Neurons with Power-of-Two Quantization Probe-Geometry Alignment: Erasing the Cross-Sequence Memorization Signature Below Chance Benchmarking local Hebbian learning rules for memory storage and prototype extraction Robust volatility updates for Hierarchical Gaussian Filtering Spiking Sequence Machines and Transformers Affinity Is Not Enough: Recovering the Free Energy Principle in Mixture-of-Experts Scalable Learning in Structured Recurrent Spiking Neural Networks without Backpropagation Geometric and dynamical analysis of attractor boundaries and storage limits in kernel Hopfield networks Attractor FCM Physical Foundation Models: Fixed hardware implementations of large-scale neural networks When Does Structure Matter in Continual Learning? Dimensionality Controls When Modularity Shapes Representational Geometry Learning to Forget: Continual Learning with Adaptive Weight Decay Causal Learning with Neural Assemblies NORACL: Neurogenesis for Oracle-free Resource-Adaptive Continual Learning Text-Utilization for Encoder-dominated Speech Recognition Models EdgeSpike: Spiking Neural Networks for Low-Power Autonomous Sensing in Edge IoT Architectures EvoTSC: Evolving Feature Learning Models for Time Series Classification via Genetic Programming Analysis and Explainability of LLMs Via Evolutionary Methods Deployment-Aligned Low-Precision Neural Architecture Search for Spaceborne Edge AI SeaEvo: Advancing Algorithm Discovery with Strategy Space Evolution Primitive Recursion without Composition: Dynamical Characterizations, from Neural Networks to Polynomial ODEs MAEO: Multiobjective Animorphic Ensemble Optimization for Scalable Large-scale Engineering Applications Necessary and sufficient conditions for universality of Kolmogorov-Arnold networks Learn&Drop: Fast Learning of CNNs based on Layer Dropping Architecture-Induced Recoverability Bias in Differentiable Symbolic Regression Collocation-based Robust Physics Informed Neural Networks for time-dependent simulations of pollution propagation under thermal inversion conditions on Spitsbergen Structure-Guided Diffusion Model for EEG-Based Visual Cognition Reconstruction
Beyond Rate Coding: Surrogate Gradients Enable Spike Timi...
Ziqiao Yu, Pengfei Sun, Danyal Akarca, Dan F. M. Goodman · 2025-07-22 · via cs.NE updates on arXiv.org

The surrogate gradient descent algorithm enabled spiking neural networks to be trained to carry out challenging sensory processing tasks, an important step in understanding how spikes contribute to neural computations. However, it is unclear the extent to which these algorithms fully explore the space of possible spiking solutions to problems. We investigated whether spiking networks trained with surrogate gradient descent can learn to make use of information that is only encoded in the timing and not the rate of spikes. We constructed synthetic datasets with a range of types of spike timing information (interspike intervals, spatio-temporal spike patterns or polychrony, and coincidence codes). We find that surrogate gradient descent training can extract all of these types of information. In more realistic speech-based datasets, both timing and rate information is present. We therefore constructed variants of these datasets in which all rate information is removed, and find that surrogate gradient descent can still perform well. We tested all networks both with and without trainable axonal delays. We find that delays can give a significant increase in performance, particularly for more challenging tasks. To determine what types of spike timing information are being used by the networks trained on the speech-based tasks, we test these networks on time-reversed spikes which perturb spatio-temporal spike patterns but leave interspike intervals and coincidence information unchanged. We find that when axonal delays are not used, networks perform well under time reversal, whereas networks trained with delays perform poorly. This suggests that spiking neural networks with delays are better able to exploit temporal structure. To facilitate further studies of temporal coding, we have released our modified speech-based datasets.