Large language models are remarkably capable, yet how computation propagates through their layers remains poorly understood. A growing line of work treats depth as discrete time and the residual stream as a dynamical system, where each layer's nonlinear update has a local linear description. However, previous analyses have relied on scalar summaries or approximate linearizations, leaving the full spectral geometry of trained LLMs unknown. We perform full Jacobian eigendecomposition across three production--scale LLMs and show that training installs a monotonic spectral gradient through depth -- from non-normal, rotation-dominated early layers to near--symmetric late layers -- together with a cumulative low-rank bottleneck that funnels perturbations into a small fraction of the residual stream's effective dimensions. Our experiments reveal that this gradient and the dimensional collapse are learned rather than architectural, and is largely dissolved when structured non-normality is removed. We further show that the topological positioning of graph communities predicts whether the Jacobian amplifies or suppresses them, with the sign of the coupling determined by the local operator type, a relationship absent at initialization. These results map a learned spectral geometry in LLMs that links perturbation propagation and compression to the network's functional topology.