We introduce E = T*H/(O+B), a dimensionless control parameter that predicts whether Mixture-of-Experts (MoE) models will develop a healthy expert ecology or collapse into dead experts. E combines four hyperparameters -- routing temperature T, routing entropy weight H, oracle weight O, and balance weight B -- into a single quantity. Through 12 controlled experiments (8 vision, 4 language) totaling over 11,000 training epochs, we establish that E >= 0.5 alone is sufficient to guarantee zero dead experts, removing the necessity for handcrafted load-balancing auxiliary losses. We validate this cross-modally on CIFAR-10, CIFAR-100, TinyImageNet-200, WikiText-2, and WikiText-103. Six additional findings emerge: (1) dead experts can resuscitate -- triggered by balance loss driving router re-exploration; (2) ortho toxicity is dataset-dependent, not universal; (3) task complexity shifts the critical E threshold; (4) model overfitting is decoupled from expert ecological health; (5) three-tier MoE spontaneously collapses into a two-tier functional structure; (6) ecological structure is temperature-invariant across a 50x range. We propose that E serves as a unified diagnostic for MoE training, analogous to the Reynolds number in fluid dynamics.